Frogmouths of the Marbled kind in Tamborine Mountain’s Escarpment

A beautiful rainforest bird, the Marbled Frogmouth (Podargus ocellatus plumiferus), is just one of the many incredible birds to make The Forest of the Tamborine Mountain Escarpment Protection Precinct their home.

The difference between Marbled Frogmouths & Tawny Frogmouths

Marbled Frogmouth (Podargus ocellatus plumiferus) in Guanaba Forest

Marbled Frogmouth

Marbled Frogmouths are very much like their Tawny counterparts (Podargus strigoides), but with some clear distinctions, including:

1. having longer tails that taper rather than flatten

2. darker yellow or orange eyes rather than yellow eyes

3. blotching underparts rather than subtle streaking

4. barred bristles appearing at the top of the beak giving them their alternative name – Plumed Frogmouths (Tawny bristles are unbarred).

But, of course, as with all Frogmouths, they freeze and become completely motionless when spotted. They do this in a bid to camouflage themselves against potential threats.

Where to find Marbled Frogmouths

Tawny Frogmouths (Podargus strigoides)

Tawny Frogmouths

Marbled Frogmouths reside in tropical and sub-tropical rainforest and montane forests.

In Australia, Marbled Frogmouths are found in North Queensland (Cape York) and from Gladstone down to the NSW border. In particular, they’re known to reside along the Canondale Ranges within Queensland’s Sunshine Coast region.

They’re also found in the Solomon Islands, Papua New Guinea and Indonesia.

Species status of Marbled Frogmouths

Marbled Frogmouth (Podargus ocellatus plumiferus) looking directly at the photographer

Marbled Frogmouth

Marbled Frogmouths are listed as “vulnerable” under Queensland’s Nature Conservation Act 1992, with the IUCN Red List recording a decline in species numbers.

Populations of Marbled Frogmouths are currently under threat by a range of human activities, including land clearing, timber harvesting and inappropriate fire regimes.

The species have a distinct need for un-logged, remnant forests for their continued survival; a vital reason to protect The Forest against inappropriate development.

More information

Sunshine Coast Council: Mary Cairncross Scenic Reserve: Marbled Frogmouth (Podargus ocellatus plumiferus)

Sunshine Coast Birds: Marbled Frogmouth & Tawny Frogmouth

Birds in Backyards: Tawny Frogmouth

IUCN Red List: Search for at-risk species around the world

Save Guanaba Facebook

Like the Save Guanaba Facebook page for regular updates on at-risk species in The Forest of Tamborine Mountain.

What are we fighting? Find out in Save Guanaba: About us

Find out more about The Forest’s precious flora and fauna

DoE: Guanaba Experience likely to negatively impact at-risk species

The Federal Department of the Environment has recently concluded the entertainment park development Guanaba Experience is likely to have a significant impact on federally protected threatened species and ecological communities.

This means, over the next few months, the Federal Department of the Environment will assess the development.

Save Guanaba Facebook

Like the Save Guanaba Facebook page for regular updates on at-risk species in The Forest of Tamborine Mountain.

What are we fighting? Find out in Save Guanaba: About us

Find out more about The Forest’s precious flora and fauna

What is the Tamborine Mountain Escarpment Protection Precinct and why bother protecting it?

Many amazing orchids of the Tamborine Mountain Escarpment

The Tamborine Mountain Escarpment Protection Precinct is not just home to an incredible array of fauna needing protection; it’s also home to an amazing variety of flora, including precious orchids.

The Forest is rich with an amazing array of epiphytic, lithophytic and terrestrial orchids. Some orchid species would have evolved to survive in the area’s unique habitats contained within gullies, on cliff faces and throughout other natural and distinct land formations.

Spotted Hyacinth Orchid

Spotted Hyacinth Orchid









The orchids in The Forest, along with a range of the other plant species, have remained mostly intact due to the area having been largely untouched by humans.

Guanaba Experience threatens their future by subjecting sensitive biodiverse areas to 50,000+ tourists per year. Orchids, which are very susceptible to subtle changes in their environment, will be exposed to additional light and soil run-off caused by vegetation clearing for tracks, trails and buildings and disturbance caused by high volumes of human activity.

Tetrabaculum Tetragonum Orchid

Tetrabaculum Tetragonum Orchid











Amazing orchids of Tamborine Mountain, many of which exist in The Forest:


Vulnerable – Bulbophyllum globuliforme (Hoop Pine Orchid)
Vulnerable – Cryptostylis hunteriana (Leafless Tongue-orchid)
Endangered – Phaius australis (Lesser Swamp-orchid)
Vulnerable – Sarcochilus hartmannii (Blue Knob Orchid)

EPIPHYTIC & LITHOPHYTIC (growing on. trees, rocks, etc.)

Bulbophyllum aurantiacum
B. crassulifolium (Wheat leafed bulbophyllum)
B. exiguum
Cymbidium madidum
C. suave
Dendrobium aemulum (Box orchid)
D. beckleri (Pencil orchid)
D. delicatum
D. gracilicaule
D. gracillimum
D. kingianum
D. linguiforme (Tongue orchid)
D. monophyllum (Lily of the valley orchid)
D. mortii
D. pugioniforme (Dagger orchid)
D. speciosum var. hilli
D. teretifolium (Bridal veil or pencil orchid)
Sarcochilus ceciliae var. albus (Fairy Bells)
S. falcatus (Orange blossom orchids)
S. fitzgeraldii (Ravine orchid)
S. hartmanii
S. ohivaceus.

TERRESTRIAL (growing in the ground)

Caladenia carnea (Pink fingers)
C. fitzgeraldii
C. patersonii (Common spider orchid)
Calanthe triplicata (Christmas orchid)
Caleana. grandflora
C. major (Flying duck or bee orchid)
Calochilus robertsonii (Bearded orchid)
Dipodium pulchellum
Diuris aurea (Double tail)
D. maculata (Spotted double-tail or leopard orchid)
D. pedunculata (Golden moth)
D. punctata
D. sulphurea
Erthrorchis cassythoides ((formerly Galeola c.) Climbing Orchid)
Geodorum neocaledonicum
Glossodia major (Wax-tip orchid)
Microtis parviflora (Slender onion orchid or babes in the wood)
Oberonia palmicola
Peristeranthus hillii
Plectorrhiza tridentata (Tangle orchid)
Prasophyllum archeri (Variable midge orchid)
Pseudovanilla foliata (formerly Galeola f.)
Pterostylis acurninata (Sharp greenhood)
P. baptistii (King greenhood)
P. concinna
P. curta
P. grandiflora (Superb or cobra greenhood)
P. longifolia
P. nutans (Nodding greenhood or Partos beak orchid)
P. obtusa
P. ophioglossa (Snakes tongue orchid)
P. reflexa (Horned orchid or dainty greenhood)
Rhinerrhiza divitzfloria
Thelymitra ixiodes (Spotted sun-orchid)

A point of interest

Renowned Australian poet Judith Wright, who was called “the conscience of the nation” for her commitment to Aboriginal Australian land rights and the nation’s natural environment, lived in a house on Tamborine Mountain called Calanthe.

The title came from the Calanthe Triplicata, a rare white orchid that flowers around December / January.

Find out more about Save Guanaba

Visit the Save Guanaba Facebook page

Tropilis Radiata Orchid

Tropilis Radiata Orchid

Magnificent Glossy Black-Cockatoos of Tamborine Mountain

The Glossy Black-Cockatoo loves the She-oaks of The Forest in the Tamborine Mountain Escarpment Protection Precinct; a good thing considering they’re such picky eaters and prefer so few plants.

Physical features of the Glossy Black-CockatooGlossy Black-Cockatoo in Guanaba Forest

Glossies are the smallest of Australia’s five black-cockatoos and are often confused with the Red-tailed Black Cockatoo (which has more white dots spread across their entire body).

Glossies have a splash of red on the underside of their tails. Adult females have some yellow splashes around their head and neck, while the red on the tails of the adult males tend to be a very bright red.

Threats to the Glossy Black-Cockatoo

Glossy Black-Cockatoos are heavily reliant on a diet of Allocasuarinas for their survival. This very specific requirement may have contributed to their dwindling numbers, as land developments and farming have led to a loss of Allocasuarina tree species. Additionally, land clearing and more frequent and intense fires have led to not only a loss of food, but also a loss of suitable nesting tree hollows. This has further placed Glossy numbers at risk.

Species status of the Glossy Black-Cockatoo

In Queensland, the Calyptorhynchus lathami lathami, which can be found in the south-east corner of the State, north and eastern NSW and a little in Victoria, is considered vulnerable under Queensland’s Nature Conservation Act (NCA). It’s also considered vulnerable in NSW and threatened in Victoria.

What Glossy Black-Cockatoos eat

As mentioned, Glossies have a very specific diet, with the Black She-oak (Allocasuarina littoralis) and the Forest She-oak (Allocasuarina torulosa) being their preferred food. They also have a tendency of returning to the same food tree time and again, ignoring trees of the same kind around them.

The Tamborine Mountain Escarpment Protection Precinct is home to both the Black and Forest She-oaks and supports a small population of Glossy Black Cockatoos (including the two pictured – a father and daughter).

How Glossy Black-Cockatoos live

Glossies are tree hollow dwellers, liking large trees with decently sized hollows for breeding and general habitat. They form monogamous pairs to breed every two years, laying a single egg that they incubate for up to 90 days.

The Glossy Black-Cockatoo count

In the October 2014, the Glossy Black Conservancy held its annual birding day to count Glossy numbers. Observers recorded 4 birds in the Scenic Rim. These results were markedly lower than those from the previous year, where observers recorded 55 birds. (Note: The drop in the number of recorded birds may be the result of a decline in spotters from 40 in 2013 to 8 in 2014.

The need to protect remnant forest to retain both food and shelter for Glossy Black-Cockatoos is essential for the survival of these beautiful birds. The Forest of the Tamborine Mountain Escarpment Protection Precinct is one such place able to sustain Glossies, which is one of the many reasons why we’re fighting for its protection against an out-of-place development.

More information:

Terrific online resource dedicated to Glossies: Glossy Black Conservancy

Seeking volunteers to monitor Glossy Black-Cockatoo numbers

Birds in Backyards: Glossy Black-Cockatoo

NSW Government Office of Environment & Heritage: Glossy black-cockatoo

Find out about this out-of-place development

See our Save Guanaba Facebook page for more about our wildlife

Lantana – manage it carefully – a lot of wildlife use it as habitat

We all recognise the massive negative impact Lantana continues to have on native flora and fauna. But there are many things to consider when managing this invasive weed, including the important habitat it provides some Australian fauna.

While Lantana threatens some 158 native animal species, it benefits another 142 native animal species. These species use the weed as habitat when relevant native flora is absent.

The types of animal species that benefit from Lantana include: mammals (27), birds (90), reptiles (12), amphibians (3) and invertebrates (10).

Rough-scaled Snake in Guanaba

Rough-scaled Snake

Some of the specific animals are: Eastern Spinebills, Fairy-wrens (Variegated, Red-backed and Superb), Rough-scaled snakes and Bandicoots.

Because of the variety and number of Australian fauna making significant use of Lantana as habitat, projects aiming to restore heavily affected areas need to do so carefully and gradually. This helps minimise impact on dependant fauna and ensures they have continued support while the change takes place.

Native plants that can replace Lantana camara
A few plants native to Tamborine Mountain will help replace the structure and function of Lantana, including:

1. Native Mulberry (or White Nettle) (Pipturus argenteus)
2. Rose-leaved Bramble (Rubus rosifolius)
3. Pink-flower Native Raspberry (Rubus parvifolius)
4. Barbwire Vine (Smilax australis).

For more information:

Tamborine Mountain Landcare
Recognising the unique beauty of Tamborine Mountain. Their mission is to protect the Mountain’s natural heritage by maintaining and enhancing the environment and its biodiversity.

Variegated Fairy-wren

Variegated Fairy-wren

All about Tamborine Mountain Landcare
The Piccabeen Bookshop – Find some great resources, including the book “Tamborine Mountain Flora and Fauna”.

Wilsons Creek Huonbrook Landcare
A group of landholders who have made a commitment to repairing and enhancing the natural landscape in the Wilsons Creek, Huonbrook and Wanganui valleys within the Byron Bay Hinterland of northern NSW.

All about weeds in northern NSW (which also affects South East Queensland)

National Lantana Management Group
Weeds of National Significance (WoNS): A joint initiatives of the States, Territories and the Australian Government. The WoNS program has now become the responsibility of Australia’s States and Territories, who’ll manage the ongoing delivery of the Lantana Strategic Plan 2012-2017.

National Lantana Management Group website
The Lantana profile (PDF)

Office of Environment & Heritage NSW
Fact sheets on Lantana, managing the impact of Lantana, the Lantana key threatening process and more.

Lantana information section on the website of Environment & Heritage NSW

Find out about this out-of-place development

See our Save Guanaba Facebook page for more about our wildlife

Guanaba developer wants to increase noise pollution levels

The Guanaba Experience developer now seeks to negotiate two “conditions” Council imposed on the entertainment park; the negotiations centering on noise pollution and staff levels.

About noise and the development

Residents, Council and individual Councillors emphasised noise as one of the main concerns about the development.

We firmly believe the noise impact caused by the Guanaba Experience development is inappropriate for the Tamborine Mountain Escarpment Protection Precinct. Nevertheless, Council’s approval of the development in June this year allowed for additional noise over and above the current noise levels experienced in the area.

The developer’s negotiating with Council to allow for significantly higher levels of noise pollution from the Guanaba Experience entertainment park.

The technical details: Council stated noise could be 5dB above LA90 from 7am to 10pm and 3dB above LA90 from 10pm to 7am. Council specified these noise levels would be “acceptable” at “sensitive places” – meaning people living nearby.

The developer’s requested Council change its condition to suit the levels of noise pollution argued for in the acoustic report for the entertainment park development application.

Given the Council justified its approval of the Guanaba Experience development on the basis of using “conditions” to mitigate impacts, we expect Council to stick with its original position on noise impacts, rather than change to meet the developer’s request.

Noise and its effect on wildlifeFemale Red Necked Wallaby Guanaba

Since the 1970s, scientists have been discovering that human-generated noise pollution can have a significant impact on fauna, including:

1. Masking important environmental cues for communication and orientation.
2. Causing physiological changes, such as increased heart rate and stress.
3. Forcing abandonment of territories.
4. Altering and even halting reproduction.

It’s important to note the effects of noise pollution from the entertainment park on the ecological functioning of the 500 acres and surrounding area doesn’t seem to have been assessed at all.

Negotiating staffing levels

Council capped staff on site at 30 at any one time. The developer has asked for this cap to be removed.

Local bird migrations in Tamborine Mountain Escarpment

Some sedentary birds have a large enough range within a local ecology that they seem migratory. This is the case for a number of birds in The Forest of the Tamborine Mountain Escarpment Protection Precinct.

Here are just some of the birds we’ve photographed during late Autumn / early Winter, which is when they appear on the eastern side of the Mountain. They disappear from the local area once the warm weather starts up again.

1. Eastern Spinebill (Acanthorhynchus tenuirostris)Eastern Spinebill Male
The Eastern Spinebill is found east of the Great Dividing Range, from north Queensland down to the Flinders Ranges in South Australia. They love nectar from a range of flowers, including grevillea flowers. Their long downturned beak is perfect for tunnelling into tubular flowers. They will also eat insects.

They’re not shy birds. In fact, they’re incredibly curious and will sneak a peak to find out what other birds are doing in their territory (without trying to scare them off) and what humans are doing close-by.

More on the Eastern Spinebill

2. Grey Fantail (Rhipidura albiscapa)Grey Fantail
The Grey Fantail is found throughout Australia in treed habitats. They feed on a range of insects by foraging at the edge of treed areas and bushes as well as in canopies. They can be easily recognised by their fanned tail and their wispy way of flying when catching flying insects.

These are gregarious birds who will fly in and out of open areas to feed.

More on the Grey Fantail

3. Golden Whistler (Pachycephala pectoralis)
The Golden Whistler is a distinctive looking bird, with its bright yellow breast contrasting strongly with their black and white neck and face.Golden Whistler

They’re found from north-eastern Queensland, down to Tasmania and across to southern South Australia and south-western Western Australia, preferring denser forests. They love insects, spiders and other small arthropods. They also eat berries.

More on the Golden Whistler

4. Grey Goshawk (Accipiter novaehollandiae)

The Grey Goshawk comes in two colour morphs, 1. completely white and 2. white-chested with grey bars and grey head and wings. The latter can be found on Tamborine Mountain and in coastal areas in northern and eastern Australia.Grey Goshawk

The Grey Goshawk loves tall forests with a closed canopy, including rainforests. Their diet consists of birds, small mammals, reptiles and insects. The larger female is capable of preying on much larger animals when compared with their male counterpart.

More on the Grey Goshawk

5. White-eared Monarch (Carterornis leucotis)

The White-eared Monarch is a pied monarch-flycatcher. They look a little bit like the very cWhite-Eared Monarchommon Magpie Lark, but with a cross of black on their face and a main black line through their eye.

They can be found on the coastal lowlands and eastern slopes of the Great Dividing Range from Cape York to north-eastern NSW. Their natural habitat is rainforest and wet sclerophyll forest, and stay mainly in the upper canopy to find food. This makes them much harder to spot.

More on the White-eared Monarch

Great places to find out about birds:

NSW Office of Environment & Heritage

Queensland Department of Environment & Heritage Protection

Birds in Backyards

BirdLife Australia


Scenic Rim Regional Council approves out-of-place development: Tamborine Mountain Escarpment Protection Precinct

On Friday, 15 May 2015, Scenic Rim Regional Council planners recommended the entertainment park be approved.

On Tuesday, 19 May 2015, a majority of Scenic Rim Regional Councillors voted in favour of the development.

The Councillors who represent Tamborine Mountain voted against the development.

The Councillors who represent other areas of the Scenic Rim voted in favour of the development.

Councillors had a week to reconsider their vote before the full Council vote on 26 May. However, once the full Council meeting was held, the vote didn’t change. Despite representations made by Tamborine Mountain Councillors, the Council decided to accept the Planning and Development Committee meeting report and approve the development. Tamborine Mountain Councillors commented further on the development requesting additional conditions be applied, especially relating to fire management.

Save Guanaba reiterates: the development for a high-volume, high-impact entertainment park goes against everything the Tamborine Mountain Escarpment Protection Precinct stands for. And we will appeal the Council’s decision in Queensland’s Planning and Environment Court.

Find out about Tamborine Mountain’s green heart – the Escarpment Protection Precinct

One of the last significant green spaces left on the Mountain …

98-196 Guanaba Road Tamborine Mountain

Queensland’s DSDIP changes agency response after developer requests amendments

After a request from the developer, the Queensland Department of State Development, Infrastructure and PlGuanaba Gorgeanning (DSDIP) has amended its concurrence agency response to the entertainment park development application targeting Guanaba Forest.

Late last year, the developer made representations to DSDIP seeking changes to the Department’s original response to the development application.

The State has amended its response following these representations. Here are the key points to the final response from the DSDIP.

Within the State-regulated vegetation area (the bottom three-quarters of the property and containing wet scleropyll forest and rainforest), the developer can:
1. Set up new infrastructure.Sclerophyll Forest with Cycads
2. Construct vehicle tracks up to 10 metres wide (in accordance with the developer’s amended track plan).
3. Clear immature tree species to establish mountain bike trails and walking tracks.
4. Clear vegetation up to 10 metres wide for zipline towers and bridge structures (supports), and up to 5 metres wide for the actual ziplines and bridges.
5. Install climbing fixtures and fittings necessary for outdoor recreation, BBQ shelters and picnic tables where no clearing of regulated vegetation is required.

In terms of roads, the developer must:
1. Move the steep decent sign from the corner 120 metres further back along Guanaba Road.Water flows in Guanaba Forest Tamborine Mountain
2. Remove vegetation from the corner of Kaiser and Guanaba Roads.
3. Dedicate a 420 square metre strip along Guanaba Road to the Department of Transport and Main Roads.

For detailed information on the response from the DSDIP, go to the development applications section of and search for the DA: MCBd14/053.

The document is titled: “Amended concurrence agency response with conditions Department of State Development Infrastructure & Planning L3 RP181081″.

Note: The State response focuses only on vegetation and Guanaba Road issues. Scenic Rim Regional Council is responsible for the overall assessment of the development application.

Dragonflies in Guanaba Forest in the Tamborine Mountain Escarpment

The Tamborine Mountain Escarpment Protection Precinct is rich with abundant insect life. One of the heroes of the Escarpment is the dragonfly, an amazing flyer, successful hunter and a most intelligent insect.

Blue Skimmer Female

Female Blue Skimmer

Dragonflies are ancient animals, having been around long before the evolution of the dinosaurs. In fact, some 250 million years ago, a species of dragonfly measured a wingspan 70cm across.

There are 6,000 species of dragonflies and damselflies in the world, with Australia housing some 320 known species.

Dragonflies – very agile flyers
Dragonflies are agile flyers, with some flying across oceans to get to their destination. They can fly up, down, forwards, backwards, left and right. They have four different flying styles; counter-stroking, phased-stroking, synchronised-stroking and gliding, and use these different styles for different reasons, such as needing to change direction very quickly (for this, they use synchronised-stroking).

Female Fiery Skimmer

Female Fiery Skimmer

Where do they live?
They start their lives as nymphs, which is the larval stage of the insect. They may spend several years as a nymph; while the adult may live for a few days or weeks only.

Dragonflies are most often seen around water, but not always, and are found on every continent except Antarctica.

Some dragonflies live in and near running water and some in still water. But, if partial to still water, they don’t cross over into running water and visa versa.

Threats to dragonfly species
Because many species of dragonfly rely on precise water temperatures, good oxygen levels and unpolluted water to survive, they can act as good bio-indicators to water quality. Some dragonflies in NSW are endangered, because their habitats have been negatively impacted on by human activity.

Australian Emerald

Australian Emerald

Loss of habitat – chiefly wetlands – threatens dragonfly populations worldwide.

For example, in Japan, the loss of 60 per cent of the country’s wetlands has forced dragonflies out of their natural habitat and into domestic ponds and local creeks.

In Africa, their numbers have dropped dramatically, making them a focus of conservation attempts on the continent.

A beneficial predator against disease

A study reported by United Press International argues small insects, such as the dragonfly, are “essential for a healthy ecosystem”. And they can contribute to protecting humans from infectious diseases, such as lime disease and malaria.

Female Blue Skimmer

Female Blue Skimmer

The report suggests that the 20th Century reduction in biodiversity might be linked to a global increase in infectious diseases in humans.

The study undertook a range of research methods, such as field surveys, lab experiments and mathematical modelling to find out if the presence of dragonflies (and other predator insects) reduces infections in frogs caused by trematodes (parasitic flatworms).

It found that where more flatworm predators existed, fewer frog infections caused by the flatworm were found.

United Press International: Small predator diversity key to a healthy ecosystem

To be identified dragonfly

To be identified

Amazing sight

According to a report cited in the New Scientist, dragonflies have between 11 and 30 different visual opsins (light-sensitive proteins in the eyes of animals).

This means dragonflies can see beyond our red, blue, green colour combination and may have vision that sets a precedent for the entire animal kingdom.

Add this to other studies, which have found dragonflies see ultraviolet as well as red, blue, green and they can recognise polarised light reflections off water and you have yourself an insect with amazing visual capacity.

New Scientist article: Dragonfly eyes see the world in ultra-multicolor

The celebrated dragonfly

Many cultures revere the dragonfly. Native American tribe, the Navajos, use them as a symbol for water purity. And in Japan they’re seen as symbols of courage, strength and happiness.

While not celebrated in Europe, they play a big part in folklore, where the dragonfly is seen as sinister. UK-ites have referred to them as the “Devil’s darning needle” and in Portugal they’ve been referred to as the “eye-snatcher”.

Dragonflies in poetry

Male Fiery Skimmer

Male Fiery Skimmer

Dragonflies have found themselves in poetry.

One of Alfred Lord Tennyson’s earlier poems celebrated the dragonfly …

Today I saw the dragon-fly
Come from the wells where he did lie.
An inner impulse rent the veil
Of his old husk: from head to tail
Came out clear plates of sapphire mail.
He dried his wings: like gauze they grew;
Thro’ crofts and pastures wet with dew
A living flash of light he flew.

There are many great online and print resources on dragonflies. Find out more about the dragonfly, an amazing animal.

They rely heavily on the protection of their habitat, chiefly, water quality for their survival. Any change to their environment can impact on the very water they need to survive.

More information:

Buglife: Saving the small things that run the planet

PNAS article on dragonfly sight: Extraordinary diversity of visual opsin genes in dragonflies

PNAS article on predator diversity: Predator diversity, intraguild predation, and indirect effects drive parasite transmission

Find a dragonfly: Australian Dragonfly Identification Key

Brisbane Dragonflies Field Guide

Australian Museum: Dragonflies and damselflies: Order Odonata